Course Information

It is important to register for classes as soon as possible to ensure timely payment of academic fees as well as your first stipend payment. Berkeley registration for fall courses typically opens in mid July; UCSF registration typically opens in August. Check your CalCentral and UCSF student portal pages for enrollment dates. Check for emails from the administration at your home campus about how many units you must enroll in at each campus. 

Feel free to reach out to peer advisors with questions about what courses to sign up for, but the most important thing is to register for the required amount of units ASAP; you can easily add and drop courses throughout the beginning of the term. Additionally, don't worry if your desired Berkeley classes are full. Just register on the waitlist and make sure to show up for the first class sessions, as students are usually able to secure a spot in these classes even if they were full at registration.

Students are required to complete a breadth requirement (Area Requirements) and a depth requirement (Major and Minor Area). Students are to maintain a minimum GPA of 3.0. Some of the courses used to satisfy Area Requirements may also be counted toward the Major or Minor Areas, but they must be taken while enrolled as a student in the program.

  1. Area Requirements
  2. Major and Minor Requirements
  3. Exceptions
  4. Coursework Recommendations
  5. Sample Courses for Majors and Minors
  6. Examples of Major and Minor Titles

Area Requirements

Students must complete the Area Requirements listed below some time during their academic career in the program. Students may apply courses taken prior to entering the program (pending approval by their Graduate Adviser) or may be selected from appropriate offerings at Berkeley or UCSF. The Area Requirements Form must be updated annually with approving signatures to the Program Administrator on the Home Campus during the Spring Semester of the first two years.

Area Semester Units Quarter Units
Anatomy, Physiology and Biology 9 13.5
Biochemistry, and Chemistry beyond General Chemistry 3 4.5
Engineering in a traditional discipline and Computer Science 7 10.5
Mathematics (beyond linear algebra and differential equations)
and Statistics
2 3.0

Major and Minor Requirements

Students must identify a Major and a Minor field in which the student will complete 16 Semester Units/24 Quarter Units and 8 Semester Units/12 Quarter Units respectively on a graded basis. This requirement is designed for students explore an area of interest in depth.

Area Semester Units Quarter Units
Major Requirements 16 24
Minor Requirements 8 12

For help in designing your Major and Minor Areas, here are a few resources:


  • Area Requirements does NOT refer to the major and minor; if you come in with a B.S. in bioengineering, you have probably already satisfied all of the Area Requirements.
  • Students may apply courses taken prior to entering the program (pending approval by their Graduate Adviser) or may be selected from appropriate offerings at Berkeley or UCSF.
  • Some of the courses used to satisfy Area Requirements may also be counted toward the Major or Minor areas (see next section), but they must be taken while enrolled as a student in the Program
  • Students who already hold a Master’s Degree or professional degree may use courses from their prior degree program toward their Minor field with approval from the Head Graduate Adviser. In this case, the Major field must be in an area complementary to the student’s prior training.
  • Up to six (6) units of undergraduate upper division courses may be applied towards the major area of study with approval from the Head Graduate Adviser.
  • One course with the S/U option (rather than letter-graded) is acceptable for meeting the requirements of the Major Area if the student is able to present sufficient justification for the inclusion of the course and following approval from both the student’s Graduate Adviser and the Head Graduate Adviser.

Coursework Recommendations

See the Peer Advising’s compiled list of coursework recommendations. Please note that course availability changes every quarter and semester and some courses are only offered every two years. Furthermore, course numbers and term availability may vary.

Semester/Quarter [may change!]
Research field
Secondary research field
Course number [may change!]
Course name
BerkeleyFallOther - StatisticsPH245Multivariate statisticsLexin LiGood basic stats class, very practical. Great professor who takes time for questions, introduces concepts with context and immediate applications. Good introduction in using R programming, no exam and one final project.
BerkeleySpringBiomedical Imaging & InstrumentationEE290PAdvanced topics in bioelectronics - Bioelectronic implantsRikky MullerGreat class on electrically-active implanted medical devices; also learned a lot about FDA regulations for medical devices
UCSFSpringOther - StatisticsComputational Biology & BioinformaticsNursing 291Applied Stat Methods For Longitudinal & Hierarchical DataLongitudinal data analysis; hands-on experience of analyzing repeated measures data, no exams
BerkeleySpringBiomedical Imaging & InstrumentationBioE C265Principles of MRIMoriel VandsburgerMust-take for anyone doing MRI research. Not recommended otherwise as it's too specific otherwise - you're better suited to just taking BioE C261 instead. This course is a follow-up to BioE C261. Includes lab sections with MRI scanners.
UCSFFallBiomedical Imaging & InstrumentationBI201Principles of MRISlightly shorter/less intense course on MRI as compared to Berkeley course (BioE265)
BerkeleySpringOther - BiochemistryMCB100BBiochemistryIn-depth biochem course for biochem undergrad majors
BerkeleyFallOther - BiochemistryCHEM135Chemical biologyEvan MillerUpper-level undergrad course; it was a great class to fulfill chemistry area requirement; valuable for understanding fundamentals of macromolecular charge and structure. Evan Miller was an excellent professor and has served on quals committees
BerkeleySpringComputational Biology & BioinformaticsCS289aIntroduction to Machine LearningJonathan ShewchukJonathan Shewchuk was an incredible professor - lectured well and gave clear explanations (probably not as good with different professors - I attended a lecture with Prof Sahai and it was extremely confusing). Great theoretical overview of lots of machine learning topics. Great fundamentals of the math in ML. Quite a large homework load. It is a very difficult course, but I learned so much & got a good grasp of the mathematical basis behind different machine learning algorithms.
UCSFWinterTissue Engineering & Regenerative MedicineBiomechanicsBioE221Tissue MechanobiologyTamara Alliston, Val Weaver, Jeff Lotz, Sophie Dumont, Aaron Fields The professors are really excited to teach this class (so excited that they were willing to teach it in Mission Bay for us even though they're all based in Parnassus), it's a great overview of interesting topics that each professor is an expert in.
BerkeleyFallBioMEMS/NanotechnologyBioE 221BioMEMSAaron StreetsIt's a lot of work (homework most weeks, midterm, poster and final project), but you end up learning a lot about micro/nanofabrication techniques and applications!
BerkeleyFallTissue Engineering & Regenerative MedicineBiomechanicsBioE 211Cell and Tissue MechanotransductionSanjay KumarIt's a great intro to a research field that lots of people in our program work in, and you learn lots of cool things about cells!
BerkeleySpringSystems & Synthetic BiologyOther - GeneticsMCB240Advanced Genetic AnalysisMCB tag teamTeaches you how to properly apply genetic concepts to experiments well (rather than just memorizing/learning biology jargon)
BerkeleySpringOther - BiologyBiomaterialsMCB250Advanced ImmunologyNice overview of the native and adaptive immune system. "The best class I took in grad school". Emphasizes thinking about experiment design to answer scientific questions.
BerkeleyFallBiomaterialsBioMEMS/NanotechnologyBioE C250Nanomaterials in MedicinePhil MessersmithCovers many material characterization techniques. In the first 1/3 of the semester, Phil typically lectures; the later 2/3 of the semester consist of journal clubs/lit-review type classes. Good practice for writing proposals; has a mock study section which is helpful when thinking about how fellowships are reviewed.
UCSFSpringDrug delivery systems & PharmacogenomicsChemistry 219Drug Discovery MinicourseMichelle ArkinIntensive minicourse over 5 weeks worth 6 UCSF units. Brought in different experts in drug development to teach different components of drug design
UCSFSpringBiomedical Imaging & InstrumentationBiophysics 210Light MicroscopyDelaine LarsenGreat course that includes both principles and hands-on experience with various forms of microscopy. Very useful course that directly improved my research
UCSFSpringTissue Engineering & Regenerative MedicineDevelopmental an Stem Cell Biology 270Stem Cell Epigenetics MinicourseBarbara PanningVery interesting course about current topics in epigenetic regulation of stem cells. Barbara Panning was a great instructor.
BerkeleyFallSystems & Synthetic BiologyBioE247Principles of synthetic biologyAdam ArkinCovers a lot of useful synthetic biology related tools and current literature. Not an easy class since co-listed with undergrads, but you get a lot of good, detailed knowledge about the field)
BerkeleyFallSystems & Synthetic BiologyBioE248Bioenergy and sustainable chemical synthesisJohn DueberGreat introduction to the field as a whole and how you can use synthetic biology and metabolic engineering in a number of fields. Not as detailed technically, but gives a great overview of the field and brings in a lot of guest speakers from industry that is great, class is easy but requires a decent bit of writing of guest reflections).
BerkeleySpringSystems & Synthetic BiologyOther - Chemical EngineeringCHMENG 274Biomolecular engineeringDave SchafferVery detailed from the chemistry side of entropy and enthalpy of proteins and cells, it goes into a lot of very technical detail about chemical engineering stuff and biology, also not an easy class, but you learn a lot)
BerkeleySpringComputational Biology & BioinformaticsOther - GenomicsPlantBio220BGenomics and computational biologyDeutschbauer5 week 1.5 credit class part of PMB grad program but you learn a lot about sequencing and how to assemble genomes)
BerkeleySpringSystems & Synthetic BiologyOther - BiochemistryPlantBio220EMicrobial physiologyHans CarlsonYou learn alot about all different types of metabolism and how it works within cells - highly recommend the class, super easy, only two short assignments and Hans is great).
BerkeleyFallNeural Systems & Vision ScienceComputational Biology & BioinformaticsVisSci 265Intro to Neural NetworksBruno Olhausen
BerkeleySpringOther - CommunicationOther - LeadershipENG295Communications for Engineering LeadersSusan Houlihan & Thomas Fitzpatrick2 day crash course in being an effective communicator
BerkeleyFallBioMEMS/NanotechnologyEE247AIntro to MEMSGreenspunPretty solid intro to MEMS class, learned a lot, would come in handy if doing MEMS research
BerkeleyFallBiomedical Imaging & InstrumentationBioE C261Medical Imaging Signals & SystemsSteve ConollyGood introduction to fundamental imaging principles with a focus on the characteristics that make images clinically relevant. Very useful background knowledge of the most common imaging modalities (no ultrasound and optics, however) that sets the foundation for future classes. Great for ensuring you have the mathematical/signal processing chops to characterize and understand imaging system theory.
UCSFSpringBiomedical Imaging & InstrumentationOther - BiochemistryBioE 241Metabolism and Magnetic Resonance SpectroscopyJohn KurhanewitczHighly informative about MR spectroscopy and how to use it to study metabolism, with sessions in the NMR lab.
BerkeleyFallComputational Biology & BioinformaticsCOMPSCI 294-148Topics in machine learning, inverse problems, and data analysis in computation neuro and medical imaging / Deep unsupervised learning [this is a special topics course which may vary from semester to semester]Miki Lustig & Chunlei LiuIf one wants to be serious about machine learning, you should take all your CS classes at berkeley, this one is of particularly high quality. Take with a buddy, these courses are a lot of work
UCSFWinterComputational Biology & BioinformaticsBMI 203Biocomputing AlgorithmsRyan HernandezVery thorough dive into designing algorithms, great for anyone doing computational biology method development
UCSFWinterDrug delivery systems & PharmacogenomicsBPS 272AAdvanced Drug DeliveryFrank SzokaInteresting overview of drug delivery field, consists of different weekly speakers and a final project with a partner. Weekly speakers are interesting because they are typically leaders in the field, either in academia or start-up founders in industry. They provide a real, and first-hand perspective on drug delivery techniques.
BerkeleySpringComputational Biology & BioinformaticsEECS 227ATOptimization Models in EngineeringFundamentals of optimization, useful for any person adjacent to algorithms. Mathematical fundamentals surrounding any optimization problem, including but not limited to machine learning.
BerkeleyFallBioMEMS/NanotechnologyBiomedical Imaging & InstrumentationEE240AAnalog Integrated CircuitsFor if you are planning to do any silicon/wafer level design or work adjacent to it. Otherwise, still a good class to understand the limitations and design parameters around instrumentation and amplifier design. Warning: probably the heaviest work load of all the Berkeley courses I took.
BerkeleyFallBiomaterialsBioE C208Biological Performance of MaterialsKevin HealyGood recap/overview of biomaterials and their various uses
UCSFWinterOther - EpidemiologyEpidemiology of AgingGood class to back to clinical relevance of disease treatments, easy A
UCSFSpringComputational Biology & BioinformaticsOther - MicrobiologyBMS 270Genomics and NGS Applications in MicrobiologyCharles ChiuGreat interactive coding class for all stages of coders (beginner to experienced)
UCSFSpringSystems & Synthetic BiologyBP 219? Biophysics: Modularity in Biology MinicourseWendall Lim & Hana El-SamadGreat intro and exposure to modular systems and systems biology
UCSFFallTissue Engineering & Regenerative MedicineOther - BiologyDevelopment & Stem Cell Biology 257Development & Stem Cell BiologySarah Knox and Julie SneddonGreat class for intro to developmental biology and practice for proposal writing
Berkeley SpringDrug delivery systems & PharmacogenomicsOther - Public HealthPBHLTH275Current Topics in VaccinologyLee RileyGreat primer on how vaccines work at a global health scale and how they are developed/tested
UCSFFallDrug delivery systems & PharmacogenomicsPHARMGENOM245ABasic Principles of Pharmaceutical SciencesKathy Giacomini, Deanna Kroetz, Les Benet, and othersReally great primer on PK/PD and ADME topics making it a foundational pick for people interested in drug delivery/pharmaceutical engineering. ESSENTIAL for anyone interested in drug delivery, and you learn from THE experts in the field (Kathy Giacomini, Les Benet, and Deanna Krotz). Just the vocabulary you learn is so important when talking about PK/PD, and it'll make you sound like you know what you're talking about when you go on job interviews
BerkeleyFallNeural Systems & Vision ScienceOther - BiologyMCB C261Cellular and Developmental NeurobiologyHillel Adesnik & Stephen BrohawnGreat discussion based course with lots of literature reading, good option for anyone interested in neurobiology at the cellular level
UCSFSpringBiomedical Imaging & InstrumentationComputational Biology & BioinformaticsBioE 245Machine Learning for Medical ImagingSri Nagarajan, Valentina PedoiaGreat course to start off or continue in machine learning for anyone in medical imaging, class broken up into unsupervised ML and supervised ML sections, covers both theoretical and practical applications
BerkeleySpringComputational Biology & BioinformaticsCSC200APrinciples and Techniques of Data ScienceJoey Gonzalez and Ani AdhikariThis class gives a very useful overview of data science and python. The homework is very practical and useful, and there is a lot of guidance so even with very little programming experience it is not too hard to learn.
BerkeleyFallOther - BiologyBiomaterialsPBHLTH 263ImmunologySarah StanleyI recommend this course purely for Sarah Stanley as a professor! She is a very good lecturer and does a good job of teaching the main principles without getting too bogged down in the specific details.
BerkeleyFallBiomedical Imaging & InstrumentationBioE 168LPractical light microscopyDan FletcherGave a hands-on approach to teaching the basics of optics with labs that demonstrated the fundamentals very well.
BerkeleyFallOther - BiochemistryMCB C100A / CHEM C130ABiophysical Chemistry: Physical Principles and the Molecules of LifeRecommended to fulfill chemistry area requirement
BerkeleyFallOther - Medical DevicesBioEng 252Clinical Needs-Based Therapy SolutionsSyed HossainyClass with guest speakers from the medical devices field (both academia and industry), and group projects related to the speaker topics
BerkeleySpringOther - EntrepreneurshipBioEng 253Biotechnology EntrepreneurshipDavid KirnClass with guest speakers from all aspects of biotech entrepreneurship (R&D, legal, business, academia...) and a group project
BerkeleySpringOther - Data ProcessingAstron 250Python Computing for Data ScienceJosh BloomExcellent, very practical overview of using Python tools for a variety of data processing tasks. Weekly homeworks take time but it's worth it; final project uses Python to do data processing for your own research
BerkeleyFallBiomaterialsBioMEMS/NanotechnologyMecEng C217Biomimetic EngineeringRobert FullFun class where you learn about biology-inspired engineering, and do a group project proposing a new biomimetic product idea
BerkeleyFallBiomedical Imaging & InstrumentationCompSci 294Computational ImagingRen NgCovers various aspects of computational imaging and has a group project at the end. The math may seem intimidating at times, but it's OK, you can do it 🙂 Especially good if you plan to work on designing microscope- or camera-based systems

Sample Courses for Majors and Minors

Here are some example courses taken by BioE students to satisfy the major and minor requirements set forth by the program. It is meant to be used as a starting point for students who would like suggestions on courses that fit their research interests. Please note that course availability changes every quarter and semester and some courses are only offered every two years. Furthermore, course numbers and term availability may vary.

Note that there are no set major or minor focus areas that you must follow, and no set courses that you must take to satisfy a particular major or minor, which allows for maximum flexibility in taking courses that help you with your research area. The rules are detailed as follows under “Major and Minor Areas” in the Grad Student Handbook.

See the Peer Advising’s compiled list of sample courses.

Course number [may change!]
Course name
BiomaterialsUCBBioEng 211Cell and Tissue Mechanotransduction
BiomaterialsUCBBioEng 221Advanced BioMEMS and Bionanotechnology
BiomaterialsUCBBioEng 224Basic Principles of Drug Delivery
BiomaterialsUCBBioEng 290AStem Cell Technologies
BiomaterialsUCBBioEng C118Biological Performance of Materials
BiomaterialsUCBBioEng C216Macromolecular Science in Biotechnology and Medicine
BiomaterialsUCBBioEng C223Polymer Engineering
BiomaterialsUCBChem 271A-CChemical Biology
BiomaterialsUCBChmEng 295DDevelopment of Biopharmaceuticals
BiomaterialsUCBChmEng 295SIntroduction to Expermental Surface Chemistry
BiomaterialsUCBMatSci C250Nanomaterials in Medicine
BiomaterialsUCBMecEng C217Biomimetic Engineering
BiomaterialsUCBPbHlth 263Public Health Immunology
BiomaterialsUCSFBIOE 221Tissue Mechanobiology
BiomaterialsUCSFBIOE 225BTissue and Organ Biology
BiomaterialsUCSFBIOE 242Principles of Tissue Engineering
BiomaterialsUCSFBPS 245APrinciples of Pharmaceutical Sciences
BiomaterialsUCSFPSPG 245BEnzymes and Reactions
BiomechanicsUCBBioEng 102Biomechanics
BiomechanicsUCBBioEng 210Cell Mechanics and the Cytoskeleton
BiomechanicsUCBBioEng 211Cell and Tissue Mechanotransduction
BiomechanicsUCBBioEng 263Principles of Molecular and Cellular Biophotonics
BiomechanicsUCBBioEng C119Occupational Biomechanics
BiomechanicsUCBBioEng C215Mechanobiology of the Cell
BiomechanicsUCBBioEng C223Polymer Engineering
BiomechanicsUCBChem 270AAdvanced Biophysical Chemistry
BiomechanicsUCBChmEng 250Transport Processes
BiomechanicsUCBMecEng 212Heat and Mass Transfer in Biology
BiomechanicsUCBMecEng 214Structure Function Relationships
BiomechanicsUCBMecEng 224Mechanical Behavior of Engineering Materials
BiomechanicsUCBMecEng C176Orthopedic Biomechanics
BiomechanicsUCSFBIOE 221Tissue Mechanobiology
BiomechanicsUCSFBIOE 240Magnetic Resonance Imaging
Biomedical Imaging & InstrumentationUCBBioEng 110Biomedical Physiology for Engineers
Biomedical Imaging & InstrumentationUCBBioEng 115Cell Biology for Engineers
Biomedical Imaging & InstrumentationUCBBioEng 164Optics and Microscopy
Biomedical Imaging & InstrumentationUCBBioEng 165Introduction to Medical Imaging
Biomedical Imaging & InstrumentationUCBBioEng 221LBioMEMS and Bionanotechnology Laboratory
Biomedical Imaging & InstrumentationUCBBioEng 263Principles of Molecular and Cellular Biophotonics
Biomedical Imaging & InstrumentationUCBBioEng 290Biomedical Imaging Signal and Systems II
Biomedical Imaging & InstrumentationUCBBioEng C117Structural Aspects of Biomaterials
Biomedical Imaging & InstrumentationUCBBioEng C213Fluid Mechanics of Biological Systems
Biomedical Imaging & InstrumentationUCBBioEng C261Medical Imaging Signals and Systems
Biomedical Imaging & InstrumentationUCBBioEng C265Principles of Magnetic Resonance Imaging
Biomedical Imaging & InstrumentationUCBCompSci 160User Interfaces
Biomedical Imaging & InstrumentationUCBCompSci 294Computational Imaging
Biomedical Imaging & InstrumentationUCBCompSci 294Design of Health Technology
Biomedical Imaging & InstrumentationUCBCompSci 294Visualization
Biomedical Imaging & InstrumentationUCBElEng 218AIntroduction to Optical Engineering
Biomedical Imaging & InstrumentationUCBElEng 225BDigital Image Processing
Biomedical Imaging & InstrumentationUCBElEng 227ATOptimization Models in Engineering
Biomedical Imaging & InstrumentationUCBElEng 236AQuantum and Optical Electronics
Biomedical Imaging & InstrumentationUCBElEng 240AAnalog Integrated Circuits
Biomedical Imaging & InstrumentationUCBElEng 290Advanced Brain Imaging Methods
Biomedical Imaging & InstrumentationUCBElEng 290CLow-Powered Bioelectronics
Biomedical Imaging & InstrumentationUCBElEng C249AIntroduction to Embedded Systems
Biomedical Imaging & InstrumentationUCBIntegBi 131Human Anatomy
Biomedical Imaging & InstrumentationUCBMecEng 109Heat Transfer
Biomedical Imaging & InstrumentationUCBMecEng 128Computer-Aided Mechanical Design
Biomedical Imaging & InstrumentationUCBMecEng 145LTransducer Lab
Biomedical Imaging & InstrumentationUCBMecEng 212Heat and Mass Transfer in Biology
Biomedical Imaging & InstrumentationUCBMecEng 235Design of Microprocessor-Based Mechanical Systems
Biomedical Imaging & InstrumentationUCBPhysics 137AQuantum Mechanics
Biomedical Imaging & InstrumentationUCBPsych 201functional Magnetic Resonance Imaging
Biomedical Imaging & InstrumentationUCBPsych 214functional Magnetic Resonance Imaging Methods
Biomedical Imaging & InstrumentationUCSFANATY 116Brain Anatomy
Biomedical Imaging & InstrumentationUCSFANATY 116Gross Anatomy
Biomedical Imaging & InstrumentationUCSFBIOE 230CIntroduction to Molecular Imaging
Biomedical Imaging & InstrumentationUCSFBIOE 230CPhysics of Radiation Oncology
Biomedical Imaging & InstrumentationUCSFBIOE 241Metabolism and Magnetic Resonance Spectroscopy
Biomedical Imaging & InstrumentationUCSFBIOE 244Medical Imaging Processing
Biomedical Imaging & InstrumentationUCSFBIOE 245Electromagnetic Neuroimaging
Biomedical Imaging & InstrumentationUCSFBIOE 245Machine Learning Algorithms for Medical Imaging
Biomedical Imaging & InstrumentationUCSFBIOE 247Introduction to MRI Systems & Hardware
Biomedical Imaging & InstrumentationUCSFBIOE 270Translational Challenges of Medical Devices
Biomedical Imaging & InstrumentationUCSFBIOE 297MRI Programming
Biomedical Imaging & InstrumentationUCSFBIOIM 201Principles of Magnetic Resonance Imaging
Biomedical Imaging & InstrumentationUCSFBIOIM 202Physical Principles of CT, PET & SPECT Imaging
Biomedical Imaging & InstrumentationUCSFBIOIM 203Imaging Probes for Nuclear and Optical Imaging
Biomedical Imaging & InstrumentationUCSFBIOIM 204Principles of Diagnostic and Therapeutic Ultrasound
Biomedical Imaging & InstrumentationUCSFBIOIM 211MR Pulse Sequences
Biomedical Imaging & InstrumentationUCSFBIOIM 220Advanced Neurological Imaging
Biomedical Imaging & InstrumentationUCSFBIOIM 230Vascular Imaging
Biomedical Imaging & InstrumentationUCSFBIOIM 240Muscoskeletal Imaging
Biomedical Imaging & InstrumentationUCSFBIOIM 250Abdominal and Pelvic Imaging
Biomedical Imaging & InstrumentationUCSFBIOIM 260Image Processing and Analysis I
Biomedical Imaging & InstrumentationUCSFBIOIM 265Image Processing and Analysis II
Biomedical Imaging & InstrumentationUCSFBIOIM 270Cancer Imaging
Biomedical Imaging & InstrumentationUCSFBIOPHYS 201Cell Biophysics
Biomedical Imaging & InstrumentationUCSFBIOSTAT 183Statistical Analysis
Biomedical Imaging & InstrumentationUCSFBIOSTAT 208Biostatistical Methods
Biomedical Imaging & InstrumentationUCSFBMS 230Advanced Topics in Cancer Research
Biomedical Imaging & InstrumentationUCSFNEUROSCI 201CIntroduction to Systems and Behavioral Neuroscience
Biomedical Imaging & InstrumentationUCSFRAD/BIOIM 209Imaging Lab: MR, CT, PET, & SPECT
BioMEMS & NanotechnologyUCBBioEng 211Cell and Tissue Mechanotransduction
BioMEMS & NanotechnologyUCBBioEng 221Advanced BioMEMS and Bionanotechnology
BioMEMS & NanotechnologyUCBBioEng 221LBioMEMS and Bionanotechnology Laboratory
BioMEMS & NanotechnologyUCBBioEng 251Micro/Nanofluidics for Bioengineering and Lab-On-A-Chip
BioMEMS & NanotechnologyUCBBioEng 263Principles of Molecular and Cellular Biophotonics
BioMEMS & NanotechnologyUCBBioEng C118Biological Performance of Materials
BioMEMS & NanotechnologyUCBBioEng C213Fluid Mechanics of Biological Systems
BioMEMS & NanotechnologyUCBBioEng C280Introduction to Nano-Science and Engineering
BioMEMS & NanotechnologyUCBChmEng 250Transport Processes
BioMEMS & NanotechnologyUCBElEng 143Microfabrication Technology
BioMEMS & NanotechnologyUCBElEng 247Introduction to MEMS
BioMEMS & NanotechnologyUCBElEng 290PBrain-Machine Interface Systems
BioMEMS & NanotechnologyUCBElEng C245Introduction to MEMS Design
BioMEMS & NanotechnologyUCBMCellBi 230Advanced Cell Biology
BioMEMS & NanotechnologyUCBMecEng 290LIntroduction to Nanobiology
BioMEMS & NanotechnologyUCBMecEng C217Biomimetic Engineering
BioMEMS & NanotechnologyUCBNSE C203Nanoscale Fabrication
BioMEMS & NanotechnologyUCSFEPI 260Development and Approval of Drugs and Devices
Computational Biology, Bioinformatics, & GenomicsUCBBioEng 143Computational Methods
Computational Biology, Bioinformatics, & GenomicsUCBBioEng 144Protein Informatics
Computational Biology, Bioinformatics, & GenomicsUCBBioEng 231Introduction to Computational Molecular Cell Biology
Computational Biology, Bioinformatics, & GenomicsUCBBioEng 247Principles of Synthetic Biology
Computational Biology, Bioinformatics, & GenomicsUCBBioEng 290BAdvanced Topics in Bioinformatics
Computational Biology, Bioinformatics, & GenomicsUCBBioEng C218Stem Cells and Directed Organogenesis
Computational Biology, Bioinformatics, & GenomicsUCBChem 220AThermodynamics and Statistical Mechanics
Computational Biology, Bioinformatics, & GenomicsUCBChem 220BStatistical Mechanics
Computational Biology, Bioinformatics, & GenomicsUCBCompSci 281AStatistical Learning Theory
Computational Biology, Bioinformatics, & GenomicsUCBCompSci 289AIntroduction to Machine Learning
Computational Biology, Bioinformatics, & GenomicsUCBCompSci 294Deep Unsupervised Learning
Computational Biology, Bioinformatics, & GenomicsUCBCompSci 294Special Topics in Deep Learning
Computational Biology, Bioinformatics, & GenomicsUCBElEng 290SMachine Learning for Sequential Decision Making
Computational Biology, Bioinformatics, & GenomicsUCBElEng 290THigh-Dimensional Data with Low-Dimensional Models
Computational Biology, Bioinformatics, & GenomicsUCBEngin 231Mathematical Methods in Engineering
Computational Biology, Bioinformatics, & GenomicsUCBMCellBi 206Physical Biochemistry
Computational Biology, Bioinformatics, & GenomicsUCBIndEng 290Data-X
Computational Biology, Bioinformatics, & GenomicsUCBMCellBi 240Advanced Genetic Analysis
Computational Biology, Bioinformatics, & GenomicsUCBMecEng 120 Computational Biology Across Multiple Scales
Computational Biology, Bioinformatics, & GenomicsUCBPbHlth 240CComputational Biostatistics
Computational Biology, Bioinformatics, & GenomicsUCBPlantBi 200BGenomics and Computational Biology
Computational Biology, Bioinformatics, & GenomicsUCBStat 141CStatistics for Bioinformatics
Computational Biology, Bioinformatics, & GenomicsUCBStat 260Probabilistic Modeling in Genomics
Computational Biology, Bioinformatics, & GenomicsUCBStat C245CBiostatistical Methods: Computational Statistics with Applications in Biology and Medicine
Computational Biology, Bioinformatics, & GenomicsUCSFBIOE 245Machine Learning Algorithms for Medical Imaging
Computational Biology, Bioinformatics, & GenomicsUCSFBIOMEDINF 203Introduction to Biocomputing Algorithms
Computational Biology, Bioinformatics, & GenomicsUCSFBIOMEDINF 206Statistical Methods for Bioinformatics
Computational Biology, Bioinformatics, & GenomicsUCSFBMS 270Genomics and NGS Applications in Microbiology (minicourse)
Computational Biology, Bioinformatics, & GenomicsUCSFBMS 270Genomics and NGS Applications in Microbiology (minicourse)
Computational Biology, Bioinformatics, & GenomicsUCSFEPI 266Mathematical Modeling of Infectious Diseases
Computational Biology, Bioinformatics, & GenomicsUCSFPHYSIOL 220Essential Immunology and Immunopathology
Drug Delivery Systems & PharmacogenomicsUCBBioEng 224Basic Principles of Drug Delivery
Drug Delivery Systems & PharmacogenomicsUCBBioEng C216Macromolecular Science in Biotechnology and Medicine
Drug Delivery Systems & PharmacogenomicsUCBBioEng C223Polymer Engineering
Drug Delivery Systems & PharmacogenomicsUCBMatSci 251Polymer Surfaces and Interfaces
Drug Delivery Systems & PharmacogenomicsUCBMatSci C250Nanomaterials in Medicine
Drug Delivery Systems & PharmacogenomicsUCBMCellBi 150Molecular Immunology
Drug Delivery Systems & PharmacogenomicsUCSFBIOPHYS 202Biophysical Methods
Drug Delivery Systems & PharmacogenomicsUCSFCHEM 241Molecular Thermodynamics
Drug Delivery Systems & PharmacogenomicsUCSFPSPG 245A-CBasic Principles of Pharmaceutical Science
Drug Delivery Systems & PharmacogenomicsUCSFPSPG 271Advanced Pharmacokinetics
Drug Delivery Systems & PharmacogenomicsUCSFPSPG 272AAdvanced Drug Delivery
Neural Systems Engineering & Vision ScienceUCBBioEng C261Medical Imaging Signals and Systems
Neural Systems Engineering & Vision ScienceUCBCompSci 281AStatistical Learning Theory
Neural Systems Engineering & Vision ScienceUCBCompSci 289AIntroduction to Machine Learning
Neural Systems Engineering & Vision ScienceUCBCompSci 294Deep Unsupervised Learning
Neural Systems Engineering & Vision ScienceUCBCompSci 294Special Topics in Deep Learning
Neural Systems Engineering & Vision ScienceUCBElEng 129Neural and Non-Linear Information Processing
Neural Systems Engineering & Vision ScienceUCBElEng 225ADigital Signal Processing
Neural Systems Engineering & Vision ScienceUCBElEng 225DAudio Signal Processing in Humans and Machines
Neural Systems Engineering & Vision ScienceUCBElEng 290Advanced Brain Imaging Methods
Neural Systems Engineering & Vision ScienceUCBElEng 290PBrain-Machine Interface Systems
Neural Systems Engineering & Vision ScienceUCBElEng 290SMachine Learning for Sequential Decision Making
Neural Systems Engineering & Vision ScienceUCBElEng 290THigh-Dimensional Data with Low-Dimensional Models
Neural Systems Engineering & Vision ScienceUCBIntegBi 245Neuroanatomy
Neural Systems Engineering & Vision ScienceUCBMCellBi 160Neuroscience
Neural Systems Engineering & Vision ScienceUCBMCellBi 163Mammalian Neuroanatomy
Neural Systems Engineering & Vision ScienceUCBMCellBi 166Biophysical Neurobiology
Neural Systems Engineering & Vision ScienceUCBMCellBi 261Cellular and Developmental Neurobiology
Neural Systems Engineering & Vision ScienceUCBMCellBi 262Advanced Topics in Systems Neurobiology
Neural Systems Engineering & Vision ScienceUCBMCellBi C261Advanced Cell Neurobiology
Neural Systems Engineering & Vision ScienceUCBNeurosc 201ABasic Concepts in Cell and Molecular Neuroscience
Neural Systems Engineering & Vision ScienceUCBNeurosc C262Circuit and Systems Neurobiology
Neural Systems Engineering & Vision ScienceUCBVisSci 212BVisual Neurophysiology
Neural Systems Engineering & Vision ScienceUCBVisSci 212CSpatial Vision and Machine Vision
Neural Systems Engineering & Vision ScienceUCBVisSci 298Computational Neuroscience
Neural Systems Engineering & Vision ScienceUCSFBIOE 245Machine Learning Algorithms for Medical Imaging
Neural Systems Engineering & Vision ScienceUCSFNEUROSCI 201CIntroduction to Systems and Behavioral Neuroscience
Neural Systems Engineering & Vision ScienceUCSFPHYS 248Analysis of Neural and Behavioral Data
Systems & Synthetic BiologyUCBBioEng 233Genetic Devices
Systems & Synthetic BiologyUCBBioEng 235Frontiers in Microbial Systems Biology
Systems & Synthetic BiologyUCBBioEng 244Introduction to Protein Informatics
Systems & Synthetic BiologyUCBBioEng 244LIntroduction to Protein Informatics Lab
Systems & Synthetic BiologyUCBBioEng 247Principles of Synthetic Biology
Systems & Synthetic BiologyUCBBioEng 248Bioenergy and Sustainable Chemical Synthesis: Metabolic Engineering and Synthetic Biology Approaches
Systems & Synthetic BiologyUCBBioEng 263Principles of Molecular and Cellular Biophotonics
Systems & Synthetic BiologyUCBBioEng C129Protein Engineering
Systems & Synthetic BiologyUCBChmEng 274Biomolecular Engineering
Systems & Synthetic BiologyUCBCompSci 289AIntroduction to Machine Learning
Systems & Synthetic BiologyUCBElEng 222Nonlinear Systems and Controls
Systems & Synthetic BiologyUCBElEng 227Optimization Models and Applications
Systems & Synthetic BiologyUCBEngin 231Mathematical Methods in Engineering
Systems & Synthetic BiologyUCBMCellBi 148Microbial Genetics and Genomics
Systems & Synthetic BiologyUCBMCellBi C112General Microbiology
Systems & Synthetic BiologyUCBPbHlth 263Public Health Immunology
Systems & Synthetic BiologyUCBPlantBi 200BGenomics and Computational Biology
Systems & Synthetic BiologyUCBPlantBi 200BMicrobial Physiology
Systems & Synthetic BiologyUCBPlantBi 220DCell Structure and Function
Systems & Synthetic BiologyUCBPlantBi 222Biochemistry of Biofuels
Systems & Synthetic BiologyUCBStat 131AIntroduction to Probability and Statistics for Life Scientists
Systems & Synthetic BiologyUCSFBIOCHEM 210Protein Homeostasis Network in Normal and Disease States (minicourse)
Systems & Synthetic BiologyUCSFBIOMEDINF 203Introduction to Biocomputing Algorithms
Systems & Synthetic BiologyUCSFBIOMEDINF 206Statistical Methods for Bioinformatics
Systems & Synthetic BiologyUCSFBIOPHYS 201AMacromolecular Structure and Interactions
Systems & Synthetic BiologyUCSFBIOPHYS 206Complex Biological Systems
Systems & Synthetic BiologyUCSFBMS 270Genomics and NGS Applications in Microbiology (minicourse)
Systems & Synthetic BiologyUCSFBMS 270The CRISPR Revolution (minicourse)
Tissue Engineering & Regenerative MedicineUCBBioEng 113Stem Cell Technologies
Tissue Engineering & Regenerative MedicineUCBBioEng 116Cell and Tissue Engineering
Tissue Engineering & Regenerative MedicineUCBBioEng 211Cell and Tissue Mechanotransduction
Tissue Engineering & Regenerative MedicineUCBBioEng 290AStem Cell Technologies
Tissue Engineering & Regenerative MedicineUCBBioEng 290HBio-Inspired Nanoscience and Nanomaterials
Tissue Engineering & Regenerative MedicineUCBBioEng C117Structural Aspects of Biomaterials
Tissue Engineering & Regenerative MedicineUCBBioEng C118Biological Performance of Materials
Tissue Engineering & Regenerative MedicineUCBBioEng C216Macromolecular Science in Biotechnology and Medicine
Tissue Engineering & Regenerative MedicineUCBBioEng C218Stem Cells and Directed Organogenesis
Tissue Engineering & Regenerative MedicineUCBBioEng C223Polymer Engineering
Tissue Engineering & Regenerative MedicineUCBIntegBi 131Anatomy
Tissue Engineering & Regenerative MedicineUCBIntegBi 245Neuroanatomy
Tissue Engineering & Regenerative MedicineUCBMatSci C250Nanomaterials in Medicine
Tissue Engineering & Regenerative MedicineUCBMCellBi 130Cell Biology
Tissue Engineering & Regenerative MedicineUCBMCellBi 150Molecular Immunology
Tissue Engineering & Regenerative MedicineUCBMCellBi C260Introduction to Neurobiology
Tissue Engineering & Regenerative MedicineUCBMecEng 280AIntroduction to the Finite Element Method
Tissue Engineering & Regenerative MedicineUCBMecEng C176Orthopedic Biomechanics
Tissue Engineering & Regenerative MedicineUCSFBIO 183Introduction to Statistical Analysis
Tissue Engineering & Regenerative MedicineUCSFBIOE 221ECM
Tissue Engineering & Regenerative MedicineUCSFBIOE 221Tissue Mechanobiology
Tissue Engineering & Regenerative MedicineUCSFBIOE 225BTissue and Organ Biology
Tissue Engineering & Regenerative MedicineUCSFBIOE 242Principles of Tissue Engineering
Tissue Engineering & Regenerative MedicineUCSFBIOIM 204Principles of Diagnostic and Therapeutic Ultrasound
Tissue Engineering & Regenerative MedicineUCSFCELLBIO 245Cell Biology

Examples of Major and Minor Titles

We would also like to emphasize that this list of sample titles is just for inspiration; it is not an exhaustive list, and students are able to design their own major and minor. The rules for doing so are detailed as follows under “Major and Minor Areas” in the Grad Student Handbook.

See the Peer Advising’s compiled list of example titles for major and minor areas.

Examples of major/minor titles
Biomedical Circuits/Devices
BioMEMS & Medical Devices
BioMEMS for Cellular Biology
Biophysics + Computational biology
Cellular & Molecular Engineering
Chemical Engineering
Communication, Education, and Advocacy in STEM
Computational Methods
Computational Neuroscience
Computational Statistics for Medical Imaging
Data Analysis and Statistics
Data Processing in Medicine & Biology
Drug Delivery
Engineering Devices and Materials for Biology and Medicine
Machine Learning
Machine Learning for Neuroprosthetics
Medical Imaging
Medical Imaging: Instrumentation and Algorithms
Medical Imaging: Physiology and Diseases
Molecular Biology
MRI Acquisition and Analysis
Neuroscience Imaging, Device and Data Analysis
Protein Biochemistry
Public Health Applications
Single-Cell Analysis
Structural biology
Structural Biology
Synthetic Biology
Systems/Computational Biology
Tissue Engineering
Translational Bioengineering Entrepreneurship